lottini also may bear a mid-longitudinal stripe of deep red that may appear almost black

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Equi.Linn Hipster underpants Horses-store.com lottini also may bear a mid-longitudinal stripe of deep red that may appear almost black

THE ALPHEID SHRIMP OF AUSTRALIA 67 8567; 3, AM P. 10322; 2, AM P. 11401; 1, AM P. 13547; 1, AM P. 13573; 1, BAU 3; 1, BAU 4; 1, BAU 15; 1, BAU 16; 1, BAU 24; 1, BAU 27; 1, BAU 32; 2, BAU 38; 4, BAU 44; 2, BAU 47; 3, BAU 48; 1, BAU 52; 1, BAU 57; 1, )C23; 2, MM 111; 1, MM 263; 1, MM421; 2, QM W. 999; 2, VM 2; 1, WM 23-65; 1, WM 118-65; 2, 75 LlZ-S (AM P. 27918).

DIAGNOSIS: Rostrum slender, acute, reaching to end of first article of antennular peduncle.

Rostral base broadened, flattened, not carinate dorsaliy, separated from carapace by deep and narrow sulci on each side.

Orbital hoods rounded laterally, bearing acute teeth on more medial portion of hood, teeth directed slightly inward.

Antennular peduncle slender, with second article variable but usually 1.7 times as long as broad, only slightly longer than visible portion of first or third.

Stylocerite reaching to middle of second antennular article.

Scaphocerite long with narrow squamous portion reaching just beyond end of antennular peduncle and with lateral tooth reaching well past end.

Carpocerite subequal to length of scaphocerite.

Lateral spine of basicerite acute, equal to or exceeding length of stylocerite.

Third maxilliped stout, articles with a ratio: 10:3:8.

Inferior margin of both first and second article bearing spines, superior margins bearing coarse setae distally.

Large chela compressed, margins rounded, without grooves or crests, 2.5 times as long as broad.

Dactylus not strongly arcuate, in males 0.3 length of entire chela with tip acute and hooked; in females dactylus rounded at tip, only 0.2 length of entire chela.

Merus with both superior and inferointernal angles rounded, although projecting; inferointernal margin with about 5 spines.

Small chela almost as long as large chela but more slender, varying from 2.5 to 3.0 times as long as broad; fingers about equal in length to palm.

Dactylus strongly curved at tip, crossing fixed finger when closed.

Inner margin of cutting face of fixed finger with distinct lamellar ridge along entire length.

Merus similar to that of large cheliped.

Ratio of carpal articles of second leg: 10:6:6:6:5; last four articles almost as broad as long.

Ischium of third legs inermous.

Merus inermous, over 3 times as long as wide.

Carpus with both margins continued as heavy projections.

Propodus with 5-7 spines, with broadened, flattened tips.

Dactylus heavy, blunt, laterally compressed, with thick longitudinal ridge on inferior face continuing around blunt tip as a curving ridge, the blunt tip thus carrying a ridge of hard chitin similar in form to a horses hoof; portion of dactylus surrounded by ridge of soft flexible chitin.

Telson 3.3 times as long as broad at posterior end.

First pair of dorsal spines placed just anterior to middle.

Inner pair of posterior spines nearly same size as dorsal spines.

DISCUSSION: This well-known species has been figured and described many times and has been known by4 names.

Coutiere in 1899 (p. 12) placedA/aevis into synonymy toA ventrosus; Holthuis in 1958 (p. 12) reviewed the synonymy and accepted Kingsley’s (1882:113) decision thatA ventrosus was a junior synonym of A.

Lottini Guerin.

Holthuis (1957,1961) also discussed the dates of publication of Guerin’s plate bearing the name Lottini (in his 1838 description he spelled it Lottinii).

The junior author also created a synonym, Crangon latipes, based on a juvenile specimen markedly different from the adult (1953:82); this was corrected in a subsequent paper (1958:165).

Finally we have reported that the largely unused name A thetis White is a synonym of A.

Lottini (B&B, 1977:282).

This species is most easily recognized and the most surely identified in its adult form in the whole genus by the unique development of the dactyls of the walking legs; smaller 68 DORA M.


BANNER specimens found in symbiosis with living pocilloporid corals which do not have this development should be carefully compared to the description of the juvenile form (loc.

Cit.) and with the other symbiont of the same corals, Synalpheus charon (Helier) (B&B, 1975:369).

BIOLOGICAL NOTES: This species is an obligate symbiont, living exclusively in living heads of the members of the Pocilloporidae, especially Pocillopora meandrina var.

Nobilis Verrill, P.

Ligulata Dana, large colonies of P.

Damicornis (Linnaeus) (=P.

Cespitosa Dana) and some of the species of the genus Seriatopora (the last reported by Patton, 1966).

This species, together with crabs of the genus Trapezia, is found in various parts of the branching colonies, but retire deeply between the branches when disturbed.

Also in the same heads, but apparently confined to the unbranched base is the smaller Synalpheus charon (Helier).

All three symbionts are of similar colour, a mottled bright orange-red ground color with a dark red mottling.


Lottini also may bear a mid-longitudinal stripe of deep red that may appear almost black.

In most of the Australian collections the exact depth at which the specimens were collected was not often clearly indicated, but as the host corals cannot survive much intertidal exposure, most must have been subtidal, and extending down to the depth limits of the host.

The specimens range in size up to 38 mm.

AUSTRALIAN DISTRIBUTION: We have a few specimens from western Australia from near Perth to the Dampier Archipelago.

In North Australia we have specimens from Darwin and the Torres Straits, but the majority of specimens range from Princess Charlotte Bay in north Queensland to Sydney.

We also have 4 specimens from Lord Howe Island.

There were no specimens from South Australia or Tasmania in our collections.

GENERAL DISTRIBUTION: This species is one of the most widespread species of the Indo-Pacific appearing, itwould seem, wherever the host corals appear, from the head of the Red Sea to east and South Africa, through the Indian and Pacific Oceans, and even extending beyond the Eastern Pacific Barrier to the mouth of the Gulf of California.

It would be expected to occur in the Ryukyus and Southern Japan as the genus Pocillopora appears there, but it has not yet been so reported. Alpheus socialis Helier Fig. 16 Alpheus socialis Helier, 1865:106, pI. 10, fig. 1.

Thomson, 1903:436, pI. 27, figs. 6-12.

Alpheus doto White, 1847:75. (nomen nudum). Previous Australian records Haswell, 1882b:190.


Sayce, 1902:155.

Port Phillip, Victoria.

SPECIMENS EXAMINED: 1 specimen from AM 32 (AM P. 27872); 4, AM 49 (AM P. 27867); 1, AM 76 (AM P. 27845); 7, AM 93 (AM P. 27890); 6, AM 122 (AM P. 27873); 1, AM 150 (AM P. 27843); 2, AM 192 (AM P. 27846); 2, AM214 (AM P. 27933); 1 AM 220 (AM P. 27866); 1, AM 233 (AM P. 27844); 1, AM 289 (AM P. 27847); 2, AM 383 (AM P. 27876); 1, AM 388 (AM P. 27868); 1, AM 395 (AM P. 27848); 1, AM 398 (AM P. 27849); 2, AM P. 3072; 1, AM P. 5029; 1, AM P. 5711; 1, AM P. 6309; 2, AM P. 6526; 3, AM P. 6912; 1, AM P. 8438; 3, AM P. 10092; 1, AM P. 10114; 15, AM P. 11734; 1, AM P. 13546; 1, AM P. 13560; 4, AM P. 13580; 1, AM P. 27874; 2, AM P. 27877.

DIAGNOSIS: As this species is so like A.

Parasocialis sp.

Nov., described and figured I -I m >rI ‘0 — DORA M.


BANNER 27893); 5, AM P. 3127; 2, AM P. 5215; 4,AM P. 8009; 3AM p, 13561; 2,AM P. 13569; 1,AM P. 27452; 13, BAU 2; 12, BAU 8; 19, BAU 9; 1, BAU 23; 1, BAU 25; 7, BAU,26; 34, BAU 34; 2, BAU 36; 8, BAU 41; 13, BAU 45; 4, BAU 46; 8, BAU 59; 3, BAU 72; 5, BAU 73; 4, BAU 74; 13, BAU75; 6,JB1; 2,JC1; 6,JC2; 2,JC9; 2,JC13; 1,JC19; 2,JC21; 1,JC26; 3,JC27; 9,JG 6-73; 1, jG 7-73; 3, jG 10-73; 1, JG 17-73; 4, MM 72; 2, QM W1224; 3, QM W2234; 4, QM W 2240; 15, QM W 2391; 1, UQ 21; 3, UQ 23; 1, UQ 27; 1, US 106163; 1, US 106166; 2, US 123567; 6, US 123568; 1, US 123602; 20, US 123603; 1, WM 36-65; 1, WM 142-65; 7, WM 275-65.

DIAGNOSIS: Rostrum acute, triangular, varying from 1.1 to 1.7 times as long as broad reaching almost to end of first antennular article.

Orbitorostral grooves shallow and rounded.

Second antennular article usually about 2 times as long as broad and varying from 1.3-2.0 times length of first; third varying from 0.6 to equal length of first.

Stylocerite acute, reaching to end of first antennular article.

Scaphocerite with lateral tooth reaching just beyond antennular peduncle; squamous portion reaching end of antennular peduncle.

Tip of carpocerite reaching to end of lateral tooth of scaphocerite.

Large chela similar to A.

Australiensis sp.

Novo (see below p. 256).

Small chela sexually dimorphic.

Male chela balaeniceps, varying from 3.1 to 4.7 times as long as broad.

In fully mature specimens the palm usually bears sculpturing similar to that of large chela but reduced; in smaller males sculpturing is greatly reduced and may be almost entirely absent.

Female chela not balaeniceps, varying from 3.5-4.7 times as long as broad.

Sculpturing on palm varying with maturity of female, with larger specimens bearing superior indentation and inferior shoulder strong but less developed than in large males while in smaller females all sculpturing may be lacking.

Ratio of carpal articles of second legs varying as indicated: 10: (6-8): (3-4): (3-4): (4-5).

Ischium of third leg usually with movable spine.

Merus inermous, varying from 3.5-5.0 times as long as broad.

Distal margins of carpus not produced into acute processes.

Propodus usually with about 10 spines.

Dactylus simple, slightly curved.

Telson 2.3 times as long as posterior margin is wide, spines on upper surface small.

DISCUSSION: After examining topotypes of A.

Lobidens we foundA.

Crassimanus to be a junior synonym.

Then, examining collections of A.

Lobidens from the Indo-Pacific area, including Australia, we separated the species into two geographic subspecies: A.


Lobidens and A.


Polynesica (1974 :429).

The pri ncipal difference between the two rested in the sculpturing of the small chela of the mature males.

In A.

Lobidens the superior margin of the palm proximal to the dactylar articulation is notched similar to that of the large chela and opposite this on the inferior margin is a strong shoulder with the groove extending into the medial face.

In A.


Polynesica these grooves are only slight constrictions in the outlines: However, the smaller males of A.


Lobidens are similar in sculpturing to A.





Lobi dens occurs in the western Pacific and Indian Ocean, A.


Polynesica has only been collected in the central Pacific.

This same type of variability was described by us for specimens of A.


Euphrosyne from Thailand (B&B, 1966b:130).

In this paper we are describing a new species, A.

Australiensis (p. 256), as closely related to A.


Lobidens and we are questioning the separation of A.

Inopinatus Holthuis and Gottlieb from A.


Lobidens (p. 241).

In a group of 11 specimens from near Brisbane, Qld. (AM 350) the inferointernal margin of the merus of the large cheliped was inermous distally.

However, as they resembled in every other character A.


Lobidens, we interpret this as a variation. \ THE ALPHEID SHRIMP OF AUSTRALIA 255 Tiwari (1963:307) described 2 males as A.

Crassimanus which we believe are two different species.

The short male he described exactly agrees with A.

Sudara, a species we described from Thailand (1966b:153, fig. 59; see also p.372 above).

In his plates, figures 25a, d and 26b, c, d, e are figures of A.


The long male he describes is probably the trueA.


Lobidens and is represented in his plates by figu res 25b, 25c and 26f.

Fourmanoir (1958:118, fig. 5) records on some specimens as A.

Crassimanus from Nosy Be Madagascar.

He figures the large chela with the distal margin of the grooves on both the superior and inferior margins as being projected and subacute, a condition that never occurs in A.


We feel these specimens were, in all probability, Alpheus edwardsii (Audouin).

BIOLOGICAL NOTES: Barnard (1r.J50:758) remarks on the colour of his Irve specimens, Greeny-brown, olive green, or smoky-grey, anterior parts of abdominal segments often white (producing a banded appearance), with or without longitudinal stripes (a median and 2 lateral) on each segment, the lower lateral stripe runs along the lower margins of the pleura and is often edged with black, a black spot in middle of the side on segments 2 and 4; telson and uropods apically blackish; chelae greeny-orange or greeny-brown, finger and thumb of large chela orange, tips dull violet, palm with a more or less brilliant cobalt-blue patch on inner (upper) surface; other legs dull pinkish Kemp (1915:301) also remarked on the black spots on each side of the second and fourth abdominal segments.

Kemp (loc.


P.300) found in his specimens from Chilka Lake thatthe species has an adaptability to extreme salinity changes.

He found that although the species lives under rocks it does not produce an elaborate burrow, but simply makes a horizontal tunnel not more than a few inches in length.

Farrow (1971 :482), on the other hand, reports that at Aldabra the burrows in the carbonate sand are elaborate with the main tunnel lying horizontal 8-13 cm below the sand and with several sets of dichotomously branched entrance burrows reaching to the surface.

He does not give the length of the horizontal portion of the burrow, but if one estimated from an approximatley 10 cm depth in his figure 17a, the burrow is near a half metre long.

McNae (1957:361) reports in South Africa they inhabit radiating burrows to the depth of 25 cm.

Farrow, McNae and others have reported that this species lives in associaton with gobiid fish (see also discussion on p. 182).

Almost all ou r ou r speci mens were collected i ntertidally, but we do have 4 speci mens that were captured in a prawn trawl at 14 fathoms in the Gulf of Carpentaria (AM 13), presumably they had been living in the mud on the bottom.

This species is also occasionally found in the bases of dead coral heads.

Our largest specimen attained the length of 44 mm but Barnard (1950:758) reports on specimens up to 55 mm in length.

AUSTRALIAN DISTRIBUTION: Our specimens have come from all warmer parts of Australia: Houtman Abrolhos, Darwin, Gulf of Carpentaria, Torres Straits and on down the east coast to Sydney, N.S.W.

We also have 17 specimens from Lord Howe Island.

GENERAL DISTRIBUTION: This species ranges throughout the entire Indo-Pacific area from the Red Sea to Hawaii, but the subspecies does not occur in the central Pacific area.

Forest and Guinot (1956:102) reported the species under A.

Crassimanus from Tunisia; it may have reached the Mediterranean via the Suez Canal. 256 DORA M.


BANNER Alpheus australiensis sp.

Novo Fig. 79 HOLOTYPE: 33 mm male from Caloundra, Qld., collected by A.


Livingstone 14/8/22. (AM P. 6352).

ALLOTYPE: 26 mm female from the same locality as the type. (AM P. 27264).

PARATYPES: 1 specimen from AM 46 (AM P. 27205); 11, AM 126 (AM P. 27221); 2, AM 144 (AM P. 27220); 1, AM 150 (AM P. 27202); 4, AM 167 (AM P. 27222); 3, AM +16 (AM P. 27203); 1, AM 349 (AM P. 27201); 10, AM 350 (AM P. 27248); 2, AM 406 (AM P. 27204); 1, AM P. 10980; 1, AM P. 27254; 13, AM P. 27255; 16, BAU 63.

DESCRIPTION: Rostrum conical, about as long as wide at base, reaching somewhat past middle of visible part of first antennular article.

Rostral carina rounded, extending posteriorly to base of eyes.

Orbital hoods not markedly inflated with frontal margi n somewhat convex; orbitorostral grooves moderate.

Ratio of antennular articles beginning with visible part of first antennular article 10:13:10; second antennular article 1.4 times as long as broad.

Stylocerite acute, reaching to end of first antennular article.

Squamous portion of scaphocerite moderately wide, reaching end of antennular peduncle; lateral tooth a little longer, outer margin straight.

Carpocerite reaching length of third antennular article past that article.

Basicerite with acute lateral tooth.

Ratio of articles of third maxilliped: 10:4:6.

Second article bearing only long hairs on inner face.

Large chela 2.4 times as long as broad, fingers occupying the distal 0.4.

Superior saddle broad and relatively shallow, proximal shoulder usually gently rounded but at times almost abrupt, distal shoulder always gently rounded.

Medial palmar depression a well-marked triangle whose apex reaches half the distance from saddle to proximal end of palm.

Lateral palmar depression quadrangular, reaching to linea impressa.

Inferior shoulder heavy and rounded; inferior notch broadly U -shaped, continuing on lateral face of palm as a well-defined but small triangular depression with rounded apex, and on medial face as a longer, broader, but less well-defined depression.

Plunger pronounced.

Merus a little longer than broad, bearing no teeth distally on inferointernal margin.

Small chela of male 3.0 times as long as broad, dactylus balaeniceps, fingers only slightly shorter than palm, both superior and inferior margins of palm with shallow, rounded indentations proximal to fingers.

Medial side of dactylar articulation bearing acute tooth.

Superior margin of dactylus with a slight subacute carina that disappears where the crest of hairs meet on the superior surface.

Merus 1.7 times as long as broad with distal margins inermous.

Small chela of female similar to male, 3.4 times as long as broad with fringe of setae well developed on medial and lateral margins of dactylus, but not reaching beyond two-thi rds length of dactylus and not meeti ng at crest.

Merus similar to male but 2.3 times as long as broad.

Ratio of carpal articles of second leg: 10:9:3:4:4.

Ischium of third leg armed with spine.

Merus inermous 4.3 times as long as broad.

Carpus 0.5 as long as merus; superior and inferior margins slightly projected distally.

Propodus 0.7 as long as merus bearing 7 spines on inferior margin and a pair distally.

Dactylus simple, slightly curved, 0.3 as long as propodus.

Telson 2.5 times as long as posterior margin is broad.

Dorsal spines of moderate size.

DISCUSSION: It is with considerable reluctance that we are naming this as a species separate from A.

Lobidens lobidens De Haan and A.

Lobidens polynesica Banner and THE AlPHEID SHRIMP OF AUSTRALIA 257 Banner (1974:429).

We separated the nominate species from the subspecies on the basis of the small chelae of mature males which carry heavy sculpturing in the nominate species that is entirely lacking in the subspecies.

The two subspecies are geographically separated, with the nominate form being found in the far western Pacific (including Australia) to the Red Sea and A.


Polynesica being confined to the archipelagoes of the central Pacific.

The central Pacific subspecies never reaches the large size at maturity attained by the western subspecies. A.

Australiensis can be firmly separated from the two subspecies of A.

Lobidens only by the characteristics of the small chelae of the males and females.

Like A.


Polynesica, the males of mature size lack the sculpturing on the small chela, and unlike both subspecie·s the chelae of the females show balaeniceps development.

These and other characteristics of less reliability are set fourth in Table 6. — Articles of third maxilliped with ratio: 10:3:7.

Inferior face of second article beset with fine setae.

Large chela compressed, 204 times as long as broad, fingers occupying the distal 004.

Plunger of dactylus of moderate development.

Superior saddle well defined, proximal shoulder overhanging, but not acute, and lying close to floor of saddle.

Distal shoulder of saddle prominent, gradually rounded.

Lateral palmar depression well defined, quadrangular, continued proximally to linea impressa.

Medial palmar depression well defined, a narrow triangle with apex reaching to proximal quarter of chela.

Inferior shoulder heavy, directed distally, in profile appearing as a heavy truncate lobe; inferodistal portion of lobe covered with papillae.

Inferolateral depression well defined, continuing up lateral face for 0.3 total height.

Medial face of palm bearing faint, narrow longitudinal groove from near proximal articulation to inferior notch.

Chela bearing long, forward directed setae on face near inferior margin; setae continuing to region of pollex.

Hirsute section of chela slightly papillose.

Merus almost 2.0 times as long as broad; superodistal margin obtuse, inferointernal margin armed with strong acute tooth subterminally and setae proximally.

Dactylus of small chela of male balaeniceps, chela almost 4.0 times as long as broad, fingers 0.6 of total length.

Superior surface of palm not rounded distally but appearing as a triangular flattened area demarked laterally by slight rounded ridge and medially b1 a low rounded crest that terminates before dactylar articulation; this development seen only in larger males.

Medial face with long setae and low papillae; bearing slight tooth at dactylar articulation; lateral surface nearly smooth.

Both fingers with dense rows of setae on margins of oppositive faces that cross at midpoint; setiferous crests of dactylus joining across superior margin near distal end.

Dactylus bearing low, thin tooth on cutting surface near dactylar articulation.

Inner face of chela bearing long fine setae, increasing towards proximal end, setae directed distally.

Merus inermous, 2.2 times as long as broad.

Dactylus of small chela of female not balaeniceps.

Fingers 1.6 times as long as palm with short fine setae on oppositive su rfaces that cross in the middle, inner face also beset with long, distally-directed setae similar to those of male chela.

Surface of chela almost smooth.

Merus 2.7 times as long as broad, inferointernal margin inermous but bearing a few setae along its enti re margi n.

Carpal articles of second legs with ratio: 10:5:2:2:3.

Ischium of third leg with spine.

Merus inermous, 4.8 times as long as broad.

Carpus 004 as long as merus, bearing a few setae distally and without extension of distal angles.

Propodus 0.7 as long as merus, bearing approximately 14 spines, more or less in pairs, along inferior border.

Dactylus simple, slender, 0.4 as long as propodus. Telson almosttwice as long as broad at posterior margin.

Anterior margin 1.4 times as broad as posterior margin.

Spines on upper surface prominent with anterior pair arising slightly anterior to middle.

Spines on posterolateral angles small, posterior margin slightly arcuate.

DISCUSSION: This species more closely resemblesAlpheus pacificus Dana than any other species in the Edwardsii Group.

It differs in the following characters: (1) A.

Pacificus does not have a balaeniceps dactylus in the male small chela and the palm does not bear a shoulder on the medial face. (2) In A.

Pacificus the surface of the inferior shoulder of the large chela is smooth while in this species it is papillose. (3) The plunger of the dactylus on the large chela is much longer in A.

Pacificus than in this species. (4) The inferodistal margin of the merus of the large chela of A.

Pacificus is inermous while in this species it carries a pronounced subterminal tooth. THE ALPHEID SHRIMP OF AUSTRALIA 263 This species is also related to other members of the Edwardsii Group in which the small chela of the male is balaeniceps and the female is not; the male chela is less than 5 times as long as broad; the rostrum is riot flattened above; the depression on the superointernal surface of the large chela is triangular instead of U -shaped; and the merus of the large cheliped is armed distally on the inferior margin.

These species include A.

Edwardsii (Audouin), A.

Chiragricus Milne-Edwards, A.

Lobidens lobidens De Haan, A.

Pareuchirus pareuchirus Coutiere, A.

Leptochirus Coutiere, and A.

Leptochiroides De Man.

It can be separated from all of these by the papillose shoulder on the inferior margin in the large chela, the papillose palm of the small chela in the male and the swelling near the superior margin proximal to the dactylar articulation.

This species is not subject to a great deal of variation.

We have found that the ratio of the fi rst two articles ofthe second leg varies from 10:5 to 10:8.

In the young specimens the papillae of the chelae are not as numerous and also in the smaller male specimens the swelling on the medial face near the superior margin of the small chela is scarcely discernible.

The species of Alpheus that Hutchings and Recher (1974) placed under the designation Alpheus species Band C (table 2, 6) are A.


The name papillosus refers to the papillae on the inferior shoulderof the large chela.

The holotype and allotype will be placed in the Australian Museum, Sydney, N.S.W.

The paratypes will be returned to the institutions that loaned them to us.

BIOLOGICAL NOTES: The habitat of this species is similar to that of A.

Pacificus, largely intertidal, under stones.

However, it has been dredged as deep as 10 fathoms and was frequently taken in the prawn trawls of Moreton Bay.

A colour note in the collection from US 106167 states Abdomen with white stripe down side.

Reddish to grey-green and white. We have specimens ranging from 22-40 mm.

AUSTRALIAN DISTRIBUTION: The specimens on the west coast came from Cockburn Sound and Exmouth Gulf.

Four of the specimens came from the Gulf of Carpentaria.

Five of the specimens came from near Kangaroo Is., South Australia.

The rest of the specimens ranged on the east coast from Cairns, Qld.

To Careel Bay, N.S.W.

At present, the species is known only from Australian waters.

Alpheus bisincisus De Haan Fig. 81 Alpheus bisincisus De Haan, 1850:179, pI. 45, fig. 3 (in text asA.

Avarus Fabricius, on plate as A.


Pearson, 1911 :182.

Tiwari, 1963:304, fig. 23.

Alpheus bisincisus malensis Coutiere, 1905a:910, pI. 86, fig. 48.

Alpheus bisincisus stylirostris Coutiere, 1905a:911, pI. 86, fig. 49.

Alpheus bisincisus variabilis De Man, 1909a:109; 1911 :406, fig. 95 a-e. SPECIMENS EXAMINED: 1 specimen from AM E. 3159; 2, AM P. 7050; 2, AM P. 7711; 1, UQ 36; 1, WM 93-65; 1, WM 144-65; 1, WM 185-65.

DIAGNOSIS: Rostrum reaching almost to end of first antennular article; varying from 1.8-3.0 times as long as broad at base.

Rostrum flattened dorsally with margins overhanging deep orbitorostral grooves and disappearing well posterior to eyes.

Orbitorostral margin varying from slightly to abruptly concave.

Visible part of first antennular article a little longer than third article, second article 1.5 times longer than 264 DORA M.


BANNER a d ~) I ‘ /—,

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    Horses-Store.com - lottini also may bear a mid-longitudinal stripe of deep red that may appear almost black